Histone 3D structures

3D Structures of histone

Updated on 14-September-2025

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• Histones
  • 2xd7, 1zr3, 2fxk, 1zr5 – hH2A macro domain – human
    
  • 6hq1 – hH1.0 - NMR
    
  • 4cay – hH2A +H2B
    
  • 1yd9 – H2A macro domain – rat
    
  • 1ghc – cH1 – chicken – NMR
    
  • 1hst – cH5
    
  • 2xql, 3c9k - cH3.2 + H4 + H2A + H2B – Cryo EM
    
  • 6k01, 6w4l - XlH2A/H2B - Xenopus laevis
    
  • 7pj1 – DmH2A + H2B – Drosophila melanogaster - NMR
    
  • 6k03 - CeH2A + H2B – Caenorhabditis elegans
    
  • 1yqa – yH1 (mutant) – yeast – NMR
    
  • 1uss, 1ust, 1uhm – yH1 globular domain – NMR
    
  • 1ku5 – HPHA – Pyrococcus horikoshii
    
  • 1b67, 1b6w, 1a7w, 1hta – MfHMFA – Methanococcus fervidus
    
  • 1bfm – MfHB - NMR
    
  • 1f1e – H – Methanopyrus kandleri
    
• Histones complex with protein
  • 6n89, 6n88 – hH1.0 + importin – Cryo EM
    
  • 6h8p – hH1 + histone-lysine methyltransferase
    
  • 3r45 - hH4 + H3-like centromeric protein + Holliday junction recognition protein
    
  • 3iid, 3iif – hH2A macro domain + ADP-ribose
    
  • 5vey - hH2A + H2B + polyUb
    
  • 5fug, 9inc - hH2A + H2B + YL-1
    
  • 4nft - hH2A + H2B + ANP32E
    
  • 6kbb - hH2A + H2B + SWR-complex protein
    
  • 7vcl – hH2A + H2B + BKRF4
    
  • 8i17 – hH2A + H2B + SPT16
    
  • 8yjf – hH2A + H2B + H3.1+ H4 + SPT16 + MCM2
    
  • 7qxb, 7trf – hH2A + H2B + telomerase reverse transcriptase + RNA – Cryo EM
    
  • 7qxa – hH2A + H2B + telomerase reverse transcriptase + adrenocortical dysplasia protein + RNA – Cryo EM
    
  • 6min, 6mio, 6miq – hH3 + TFIID
    
  • 6mil, 6mim – hH3 + AF9
    
  • 8ge0 – hH3 + JADE1
    
  • 8ihn – hH3 + Rpd3 + SIN3 + EAF3 + RCO1 - Cryo EM
    
  • 8z50 – hH3.1 + H4 + ASF1A
    
  • 7v6q – hH3.1 + H4 + NASP + ASF1A
    
  • 8iqf, 8iqg – hH3.1 + H4 + CAF1 + RBBP4
    
  • 6vil – hH3.1 + BAHCC1
    
  • 5bo0, 5bnx – hH3.2 + H4 + ASF1B + MCM2
    
  • 5ja4, 5bnv – hH3.3 + H4 + TONSL + MCM2
    
  • 7cj0 – hH3.3 + H4 + HDJC9 + MCM2
    
  • 7vcq – hH3.3 + H4 + ASF1B + BKRF4
    
  • 6hbo – hH3.3 + histone-lysine methyltransferase - NMR
    
  • 7ciz – hH3.3 + H4 + MCM2
    
  • 8vld – hH3.5 + histone-lysine methyltransferase
    
  • 9im4 – hH3.5 + AF-9
    
  • 9c0o – hH3.5 + CFP1
    
  • 8uhl, 8uk5 – hH4S1 + ATAD2B
    
  • 3nqj, 3nqu - hH4 + H3-like centromeric protein
    
  • 6mlc - hH4 + histone-lysine methyltransferase
    
  • 5g2e, 6k00, 8xaa - XlH2A + H2B + NAP1
    
  • 6n1z - XlH2A + H2B + importin
    
  • 8zvy - XlH2A + H2B + synuclein peptide
    
  • 7unk - XlH3 + H4 + importin-4 + ASF1
    
  • 2io5 - XlH3.1 + H4 + anti-silencing protein
    
  • 5bsa, 5bs7 – XlH3.2 + H4 + SPT2
    
  • 6o22 - XlH3.2 + H4 + RTT109 + ASF1
    
  • 4eo5 - XlH3.2 (mutant) + H4 (mutant) + ASF1
    
  • 4ld9 - XlH3.2 + H4 + H2A + H2B + widom sequence reverse + SIR3
    
  • 9b2u – XlH3.2 (mutant) + Haspin – Cryo EM
    
  • 4h9n, 4h9o, 4h9p, 4h9q, 4h9r, 4h9s, 4hga - XlH3.3 (mutant) + H4 + death domain-associated protein
    
  • 5kdm - XlH3.3 + H4 + death domain-associated protein + tegument protein
    
  • 3qzs, 3qzt – XlH4 + nucleosome-remodeling factor subunit BPTF
    
  • 6o7g - XlH4 + histone-lysine methyltransferase – NMR
    
  • 6k09, 6k0c - CeH2A + H2B + NAP1
    
  • 4uuz – DmH3 + H4 + DNA replication licensing factor
    
  • 6m2m – AtH2A + H2B + OSCHZ1 – Arabidopsis thaliana
    
  • 7c7x – AtH2A + H2B + NAP1
    
  • 9b3f – yH2A + H2B + NAP1 – Cryo EM
    
  • 9b31 – yH2A + H2B + NAP1 + KAP114 – Cryo EM
    
  • 9b3i – yH2A + H2B + NAP1 + KAP114 + GTP-binding nuclear protein – Cryo EM
    
  • 8f0x – yH2A + H2B + importin
    
  • 8f1e – yH2A + H2B + Gsp1 + importin – Cryo EM
    
  • 2l5a - yH4/H3-like centromeric protein/SCM3
    
  • 2jss - yH2A/H2B + CHZ1 – NMR
    
  • 4wnn – yH32A + H2B + SPT2
    
  • 2hue - yH3 + H4 + anti-silencing protein
  • 7xay - yH3 + H4 + cytochrome b562 + ASF1
    
  • 9isz, 9it4 – H3.1/H3.2 + histone-lysine methyltransferase – fission yeast
    
  • 2yfv, 2yfw - KlH4 + H3-like centromeric protein + SCM3 – Kluyveromyces lactis
    
  • 6uph - KlH3-like centromeric protein + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 1pfb – H3 + polycomb protein – Strongylocentrus purpuratus
    
• Human nucleosome core + DNA
  • 3av1, 2av2, 3av2, 3waa, 3wa9, 3w99, 3w98, 3w97, 3w96, 5x7x, 3a6n, 3afa, 2cv5, 5jrg, 5gxq, 5gt3, 5gt0, 5gsu, 5gse, 5cpk, 5cpj, 5cpi, 5b33, 5b32, 5b31, 5b2j, 5b2i, 5b24, 5b0z, 5b0y, 5ay8, 5avc, 5avb, 5av9, 5av8, 5av6, 5av5, 5z5t, 4ym6, 4ym5, 5y0c, 5onw, 5ong, 5omx, 6hkt, 6jou, 6jr0, 6jr1, 6k1i, 6k1j, 6j1k, 6ke9, 6l9h, 6le9, 6ipu, 6iq4, 6jxd, 6kvd, 6ler, 6m3v, 6m44, 6v2k, 8qkt, 8yti – hH3.1 + H4 + H2A + H2B in nucleosome
    
  • 6l49, 6l4a, 6l9z, 6m4d, 6m4g, 6m4h, 6r0c, 6r25, 6r93, 6r94, 6t79, 6t93, 7nl0 7u51, 7u52, 7u53, 7v90, 7v9c, 7v9j, 7v9k, 7v9s, 7va4, 7v96, 7xd1, 7xzy, 7y00, 8aag, 8guk, 8ihl, 8jbx, 8jcc, 8jcd, 8jla, 8jlb, 8jld, 8kb5, 8ke0, 8ooa, 8ox0, 8rgm, 8vlr, 8vmn, 8vo0, 8vws, 8vwu, 8y3c, 8y3d, 8y3e, 8y3f, 8ybj, 8zjt, 9be5, 9be6, 9d3n, 9d3o, 9d3p, 9d3q, 9d3r, 9d3s, 9d3t. 9dwf, 9dwi, 9dwj, 9dwk, 9dwl, 9ge4, 9geo, 9gfb – hHistones in nucleosome – Cryo EM
    
  • 5z30 – hH3.1 + H4 + H2A (mutant) + H2B in nucleosome
    
  • 5y0d – hH3.1 + H4 + H2A + H2B (mutant) in nucleosome
    
  • 4qlc – hH3 + H4 + H2A + H2B + H5 in nucleosome
    
  • 3ayw, 3aze, 3azf, 3azg, 3azh, 5b40 - hH3 (mutant) + H4 + H2A + H2B in nucleosome
    
  • 3azi, 3azj, 3azk, 3azl, 3azm, 3azn - hH3.1 + H4 (mutant) + H2A + H2B in nucleosome
    
  • 8ybk - hH3.1 (mutant) + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 7lya - hH3.1 + H4 + H2B in nucleosome – Cryo EM
    
  • 6pwx - hH3.2 + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 7pet, 7peu, 7pev, 7pew, 7pex, 7pey, 7pez, 7pf0, 7pf2, 7pf3, 7pf4, 7pf5, 7pf6, 7pfa, 7pfc, 7pfd, 7pfe, 7pff, 7pft, 7pfu, 7pfv, 7pfw, 7pfx, 9cau - hH3.2 + H4 + H2A + H2B + H1.4 in nucleosome – Cryo EM
    
  • 7x57, 7x58 - hH3.1 + H4 in nucleosome – Cryo EM
    
  • 5xf5, 5xf4, 5xf3 - hH3.1 + H4 + H2A + H2B in nucleosome + Ru derivative
    
• Human nucleosome core + DNA complex with protein
  • 5x0y, 5x0x – hH3 + H4 + H2A + H2B in nucleosome + SNF2
    
  • 9gxa, 9k9l – hH4 + DNA + CENP-A – Cryo EM
    
  • 3an2, 4x23 – hH4 + H2A + H2B in nucleosome + CENP-A
    
  • 7uv9, 9nqu – hHistones in nucleosome + lysine-specific histone demethylase – Cryo EM
    
  • 6r25 – hH3 + H4 + H2A + H2B in nucleosome + NPAC + lysine-specific histone demethylase – Cryo EM
    
  • 6r92, 6r91, 6r90, 6r8z, 6r8y - hH3.1 + H4 + H2A + H2B in nucleosome + DNA damage-binding protein – Cryo EM
    
  • 5e5a - hH3.2 + H4 + H2A + H2B in nucleosome + IE1
    
  • 8off - hHistones in nucleosome + BRCA1+ Ub – Cryo EM
    
  • 7e8i, 7lyb, 8grq - hH3.1 + H4 + H2B in nucleosome + BRCA1 + BARD1 + UbCH5C – Cryo EM
    
  • 8grm - hH3.1 + H4 + H2B in nucleosome + Commd3 + Ring1B + UbCH5B – Cryo EM
    
  • 7jzv - hH3.2 (mutant) + H4 + H2A + H2B in nucleosome + BRAC1 – Cryo EM
    
  • 9b3p - hH3.3 + XlH4 + mH2A + XlH2B in nucleosome – Cryo EM
    
  • 6uxw - hH3.3 + H4 + H2A + H2B in nucleosome + SWI/SNF complex – Cryo EM
    
  • 7ea8 - hH3.3 + H4 + H2A + H2B in nucleosome + SETD2 – Cryo EM
    
  • 7d1z - hH3.1 + H4 + H2A + H2B in nucleosome + SET8 – Cryo EM
    
  • 7d20 - hH4 + H2A + H2B in nucleosome + SET8 + Cenp-A – Cryo EM
    
  • 7r5r - hH4 + H2A + H2B in nucleosome + CenP-A + CenP-C – Cryo EM
    
  • 7y7i - hH4 + H2A + H2B in nucleosome + CenP-A + KNL2 – Cryo EM
    
  • 6muo, 6mup, 7bxt, 7by0 - hH4 + H2A + H2B in nucleosome + CenP-A + CenP-C + CenP-N – Cryo EM
    
  • 7pii - hH4 + H2A + H2B in nucleosome + CenP-C – Cryo EM
    
  • 7u46, 7u47 - hH4 + H2A + H2B in nucleosome + CenP-A + Cenp-N – Cryo EM
    
  • 7ywx, 7yyh - hH4 + H2A + H2B in nucleosome + CenP-A,C,H-Q,R,U – Cryo EM
    
  • 7xd0 - hH4 + H2A + H2B in nucleosome + L40 + histone domain-containing protein – Cryo EM
    
  • 6x59, 6x5a, 6xjd, 6y5d, 6y5e, 7a08, 7jo9, 7joa - hH3.3 + H4 + H2A + H2B in nucleosome + cGAS – Cryo EM
    
  • 7c0m, 7ccq, 7ccr - hH3.1 + H4 + H2A + H2B in nucleosome + cGAS – Cryo EM
    
  • 7bwd - hH3.1 + H4 + H2A + H2B in nucleosome + DOT1 + Ub – Cryo EM
    
  • 7xcr, 7xct - hH4 + H2A + H2B in nucleosome + DOT1L + Ub – Cryo EM
    
  • 7lyc - hH3.1 + H4 + H2A + H2B in nucleosome + BARD1 + Ub – Cryo EM
    
  • 7scy, 7scz - hH3.1 + H4 + H2A + H2B in nucleosome + PARP1 – Cryo EM
    
  • 7e8d - hH3.1 + H4 + H2A + H2B in nucleosome + NSD2 – Cryo EM
    
  • 6usj - hH3.1 + H4 + H2A + H2B in nucleosome + PARP2 – Cryo EM
    
  • 6t7a, 6t7b, 6t7c, 6t7d - hH3.2 + H4 + H2A + H2B in nucleosome + SOX – Cryo EM
    
  • 6t90, 6yov - hH3.2 + H4 + H2A + H2B in nucleosome + SOX + OCT4 – Cryo EM
    
  • 8sps, 8spu - hHistones in nucleosome + scFv + OCT4 – Cryo EM
    
  • 8ott - hHistones in nucleosome + Myc + MAX – Cryo EM
    
  • 8ots - hHistones in nucleosome + OCT4 + Myc + MAX – Cryo EM
    
  • 6rny - hH3.3 + H4 + H2A + H2B in nucleosome + integrase – Cryo EM
    
  • 6upk, 6upl - hH3.1 + H4 + H2A + H2B in nucleosome + FACT complex – Cryo EM
    
  • 6v92 – hH3.1 + H4 + H2A + H2B in nucleosome + RSC complex – Cryo EM
    
  • 8g57 – hH3 + H4 + H2A + H2B in nucleosome + sirtuin 6 deacylase – Cryo EM
    
  • 7u50 – hH3 + H4 + H2A + H2B in nucleosome + APE1 – Cryo EM
    
  • 7xzx, 7xzz, 9r04, 9r2m, 9r2p, 9r2q – hHistones in nucleosome + p53 – Cryo EM
    
  • 7w9v, 8hah, 8hai, 8haj, 8hak – hHistones in nucleosome + p300 – Cryo EM
    
  • 8dk5, 8evg, 8jl9, 8vfx – hHistones in nucleosome + scFv – Cryo EM
    
  • 9gmk, 9gmr – hHistones in nucleosome + SIRT7 – Cryo EM
    
  • 8jnd, 8jne, 8jnf, 8xbt, 8xbu, 8xbw – hHistones in nucleosome + Rad51 – Cryo EM
    
  • 8zjr – hHistones in nucleosome + RFX5 – Cryo EM
    
  • 8evh, 8evi, 8evj – hHistones in nucleosome + scFv + Pu.1 – Cryo EM
    
  • 8jhf, 8jhg – hHistones in nucleosome + SUV420H1 – Cryo EM
    
  • 7zi4, 8av6, 8oos, 9cb7, 9cf6, 9gfm – hHhistones in nucleosome INO80 complex – Cryo EM
    
  • 8atf – hH3.2 + H4 + H2A + H2B in nucleosome + INO80 + PAPA-1 domain-containing protein – Cryo EM
    
  • 7yqk – hH3.1 + H4 + H2A + H2B in nucleosome + polyUb + 53bp1 + TP53-binding protein – Cryo EM
    
  • 7tan – hH3.2 + H4 + H2A + H2B in nucleosome + VRK1 – Cryo EM
    
  • 7y8r - hH3 + H4 + H2A + H2B in nucleosome PBAF complex - Cryo EM
    
  • 6fml, 8oo7, 8oop – hHistones in nucleosome + RuvB + actin-related protein + IES6 - Cryo EM
    
  • 9gcg, 9gev – hHistones in nucleosome + RuvB + actin-related protein + INO80 - Cryo EM
    
  • 9d3m – hHistones in nucleosome + HMG-17 - Cryo EM
    
  • 8hag – hHistones in nucleosome + histone acetyltransferase - Cryo EM
    
  • 8vwt, 8vwv, 9eoz – hHistones in nucleosome + glycosylase/DNA lyase - Cryo EM
    
  • 8txv, 8txw, 8txx – hHistones in nucleosome + Rnf168 + polyUb - Cryo EM
    
  • 8iej – hHistones in nucleosome + Rnf20 + Rnf40 + Rad6A + polyUb - Cryo EM
    
  • 8u13 – hHistones in nucleosome + Rnf168 - Cryo EM
    
  • 8smz, 8sn0, 8sn1, 8sn2, 8sn3, 8sn4, 8sn5, 8sn6, 8sn7, 8sn8, 8sn9, 8sna, 8upf, 8x7i, 8x7j, 8x7k – hHistones in nucleosome + Rnf168 + Ub conjugating enzyme E2 + polyUb - Cryo EM
    
  • 8smw, 8smx, 8smy, 8u14 – hHistones in nucleosome + Rnf168 + Ub conjugating enzyme E2 - Cryo EM
    
  • 8gui, 8guj – hHistones in nucleosome + Bre1 - Cryo EM
    
  • 8ieg – hHistones in nucleosome + Bre1 + Rad6A + Ub - Cryo EM
    
  • 8osj, 8osk, 8osl – hHistones in nucleosome + Clock + Bmal1- Cryo EM
    
  • 8kd2, 8kd3 – hHistones in nucleosome + RPD3 + SIN3 + EAF3 + RCO1 - Cryo EM
    
  • 8q3m, 8q3x – hHistones in nucleosome + Kasposi sarcoma peptide
    
  • 8q36, 8q3e – hHistones in nucleosome + Foamy virus peptide
    
  • 8hal, 8ham, 8han – hHistones in nucleosome + CBP - Cryo EM
    
  • 9dwg, 9dwh, 9dwm – hHistones in nucleosome + DNA polymerase beta – Cryo EM
    
  • 8a3y, 8bvw, 8byq, 8bz1, 8h0v, 8h0w 8he5, 8jh2, 8jh3, 8jh4, 8xrj, 8xvs, 9egx, 9egy, 9egz, 9eh0, 9eh1, 9eh2, 9ii7, 9j0n, 9j0o, 9j0p – hHistones in nucleosome + RNA polymerase complex – Cryo EM
    
  • 8rbx – hHistones in nucleosome + RNA polymerase complex + PP2A – Cryo EM
    
  • 8gxq, 8gxs – hHistones in nucleosome + PIC-Mediator complex– Cryo EM
    
  • 8qzm – hHistones in nucleosome + DNMT3 – Cryo EM
    
  • 8u5h – hHistones in nucleosome + DNMT + Ub – Cryo EM
    
  • 8wg5 – hHistones in nucleosome + Usp16 + Ub – Cryo EM
    
  • 8ox1 – hHistones in nucleosome + TRF1 – Cryo EM
    
  • 8x15, 8x19, 8x1c, 9ca7, 9ca8, 9caa, 9cab – hHistones in nucleosome + SRCAP-C - Cryo EM
    
  • 9j8m, 9j8n, 9j8o – hHistones in nucleosome + BAF + lamin – Cryo EM
    
  • 8yny – hHistones in nucleosome + CAS9/CSN1 – Cryo EM
    
  • 8vg0 – hHistones in nucleosome + GATA4 – Cryo EM
    
  • 8vg1 – hHistones in nucleosome + GATA4 + FoxA1– Cryo EM
    
  • 8j6s, 8j6t – hHistones in nucleosome + CAF-1 – Cryo EM
    
  • 8uxq – hHistones in nucleosome + HP1 – Cryo EM
    
  • 8vx6 – hHistones in nucleosome + OGG1 – Cryo EM
    
  • 8kcy, 8kd1, 9l1x, 9l22 – hHistones in nucleosome + DEK– Cryo EM
    
  • 8hqy, 8hr1 – hHistones in nucleosome + SSX1 – Cryo EM
    
  • 9d3k - hHistones in nucleosome + Dsup – Cryo EM
    
  • 8vfy, 8vfz - hHistones in nucleosome + FoxA1 – Cryo EM
    
  • 8t9f - hHistones in nucleosome + histone-lysine methyltransferase – Cryo EM
  • 8bov, 8vnv, 8vnz - hHistones in nucleosome + PRC2 – Cryo EM
    
  • 8yv8 - hHistones in nucleosome + CDCA7 – Cryo EM
    
  • 8ol1 - hHistones in nucleosome + SPSB3 + ELONBC – Cryo EM
    
  • 8syp - hHistones in nucleosome + CX3CR1 – Cryo EM
    
• Xenopus nucleosome core + DNA
  • 6px1, 6r1t - XlH3 + H2A + H2B in nucleosome – Cryo EM
    
  • 1aoi, 1kx3, 1kx4, 1kx5, 1m18, 1m19, 1m1a, 1p34, 1p3a, 1p3b, 1p3f, 1p3g, 1p3i, 1p3k, 1p3l, 1p3m 1p3o, 1p3p, 1s32, 1zbb, 2fj7, 2nzd, 3c1b, 3c1c, 3kuy, 3lel, 3lz0, 3lz1, 3mnn, 3o62, 3reh, 3rei, 3rej, 3rek, 3rel, 3ut9, 3uta, 3utb, 4z66, 4ys3, 4xzq, 5cp6, 5dnm, 5dnn, 5f99, 5oxv, 5oy7 - XlH3.2 + H4 + H2A + H2B in nucleosome
    
  • 6wz5, 6wz9, 7kbd, 7kbe, 7m1x, 7ohc, 7xfc, 7xfi, 7xfl, 7xfn, 7xnp, 8cze, 8eu2, 8vx5, 8xjv, 9gd0 - XlH3.2 + H4 + H2A + H2B in nucleosome - Cryo EM
    
  • 7kbf - XlH3.2 + H4 + H2A + H2B + H1.8 in nucleosome - Cryo EM
    
  • 7swy - XlH3 (mutant) + H4 + H2A + H2B (mutant) n nucleosome - Cryo EM
    
  • 7ktq, 9c9x - XlH3 + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 6esf, 6esg, 6esh, 6esi, 6fq5, 6fq6, 6fq8, 6nn6, 6nog, 6nzo, 6px3, 6r1u, 6ugm - XlH3.2 + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 4xuj - XlH3.2 + H4 + H2A + H2B in nucleosome + Ru derivative
    
  • 4xu9 - XlH3.2 (mutant) + H4 + H2A + H2B in nucleosome + Pt derivative
    
  • 4xu8 - XlH3.2 + H4 + H2A + H2B (mutant) in nucleosome + Pt derivative
    
  • 2f8n - XlH3.1 + H4 + H2A + H2B + H3 + H2A macro domain in nucleosome
    
  • 1zla – XlH3 + H4 + H2A + H2B + H3 in nucleosome + antigen
  • 3b6f, 3b6g, 3lja, 3mgp, 3mgq, 3mgr, 3mgs, 4j8u, 4j8v, 4j8w, 4j8x, 4kgc, 5xf6 - XlH3.2 + H4 + H2A + H2B in nucleosome + metal
    
  • 3kxb - XlH3.2 (mutant) + H4 + H2A + H2B (mutant) in nucleosome
    
  • 3kwq - XlH3.2 (mutant) + H4 + H2A + H2B in nucleosome
    
• Xenopus nucleosome core + DNA complex with protein
  • 6pwx - XlH3.2 + H4 + H2A + H2B in nucleosome + RBBP5 - Cryo EM
    
  • 6pww - XlH3.2 + H4 + H2A + H2B in nucleosome + RBBP5 + WDR5 - Cryo EM
    
  • 7mbm, 7mbn - XlH3 + H4 + H2A + H2B in nucleosome + RBBP5 + WDR5 + ALL-1 + ASH2-like protein - Cryo EM
    
  • 7ud5 - XlH3 + H4 + H2A + H2B (mutant) in nucleosome + RBBP5 + WDR5 + cDNA + ASH2-like protein + DPY-30 + polyUb- Cryo EM
    
  • 8du4 - XlH3 + H4 + H2A + H2B (mutant) in nucleosome + RBBP5 + WDR5 + polyUb- Cryo EM
    
  • 7vdt - XlH3 + H4 + H2A + H2B in nucleosome + BRG1 - Cryo EM
    
  • 7vdv - XlH3 + H4 + H2A + H2B in nucleosome PBAF complex - Cryo EM
    
  • 6om3 - XlHistones in nucleosome + origin recognition complex
    
  • 8siy - XlHistones in nucleosome + origin recognition complex – Cryo EM
    
  • 8g8g - XlHistones in nucleosome + OCT4 – Cryo EM
    
  • 8ruq - XlHistones in nucleosome + borealin – Cryo EM
    
  • 8rup - XlHistones in nucleosome + borealin + inner centromere protein + surviving – Cryo EM
    
  • 8t3t, 8t3w, 8t3y - XlHistones in nucleosome + bre1 – Cryo EM
    
  • 6vyp - XlH3 (mutant) + H4 + H2A (mutant) + H2B (mutant) in nucleosome + LSD1 + COREST
    
  • 6ryr, 6ryu, 7otq, 7tn2, 9ear, 9gd1, 9gd2, 9gd3, 9nh8 – XlHistones in nucleosome + CHD1 – Cryo EM
    
  • 7cro - XlH3 + H4 + H2A + H2B in nucleosome + NSD2 – Cryo EM
    
  • 8f86, 8of4, 9eil – XlH3 + H4 + H2A + H2B in nucleosome + sirtuin 6 deacylase – Cryo EM
    
  • 7crp, 7crq, 7crr – XlHistones in nucleosome + NSD3 – Cryo EM
    
  • 9gen, 9gep, 9geq, 9ger, 9ihd, 9ihe, 9ihf – hHistones in nucleosome + myeloperoxidase – Cryo EM
    
  • 8skz – hHistones in nucleosome + DDM1/HELLS – Cryo EM
    
  • 7at8 - XlH3 + H4 + H2A + H2B in nucleosome + EZH2 + SUZ12 – Cryo EM
    
  • 6vz4 - XlH3 + H4 + H2A + H2B in nucleosome + STH1 + ARP7 + ARP9 + RTT – Cryo EM
    
  • 6kiu, 6kiv, 6kiw, 6kix, 6kiz, 6pwv, 6w5i, 6w5m, 6w5n - XlH3.2 + H4 + H2A + H2B in nucleosome + MLL1 core complex – Cryo EM
    
  • 7ohb - XlH3.2 + H4 + H2A + H2B in nucleosome + TBP – Cryo EM
    
  • 7oh9, 7oha - XlH3.2 + H4 + H2A + H2B in nucleosome + TBP + TFIIA – Cryo EM
    
  • 7ea5 - XlH3 (mutant) + H4 + H2A + H2B in nucleosome + SET2 – Cryo EM
    
  • 6s01, 7pgv, 7ph5, 7ph6 - XlH3.2 + H4 + H2A + H2B in nucleosome + PC4 and SFRS1-interacting protein – Cryo EM
    
  • 6iy3, 6iy2, 8v4y, 8v6v, 8v7l, 9e1l, 9e1m, 9e1n, 9e1o, 9e1p, 9e1q, 9e1r, 9e1u, 9e1v, 9e1w, 9e1x, 9e1y - XlHistones in nucleosome + Snf2h – Cryo EM
    
  • 7eg6 - XlH3.2 + H4 + H2A + H2B in nucleosome + SNF5 – Cryo EM
    
  • 7egp - XlH3.2 + H4 + H2A + H2B in nucleosome + SWI/SNF – Cryo EM
    
  • 6ltj - XlH3.3 + H4 + H2A + H2B in nucleosome + BAF complex – Cryo EM
    
  • 5o9g, 9n6h, 9n6i, 9n6k - XlHistones in nucleosome + CHD1 – Cryo EM
    
  • 9b2s, 9b2t - XlHistones in nucleosome + Haspin – Cryo EM
    
  • 8cbq - XlHistones in nucleosome + p75 – Cryo EM
    
  • 9d3l - XlHistones in nucleosome + Dsup – Cryo EM
    
  • 8cbn - XlHistones in nucleosome + LEDGF – Cryo EM
    
  • 6x0n - XlH3.2 + H4 + H2A + H2B in nucleosome + PARP2 + HPF1 – Cryo EM
    
  • 7nkx – XlH3 + H4 + H2A + H2B in nucleosome + CHD1 + RNA polymerase – Cryo EM
    
  • 7nky – XlH3 + H4 + H2A + H2B in nucleosome + SPT + FACT + RNA polymerase – Cryo EM
    
  • 6jm9 - XlH3.2 + H4 + H2A + H2B in nucleosome + DOT1 – Cryo EM
    
  • 8g6g, 8g6h, 8g6s, 8v27, 8v28 - XlHistones in nucleosome + Ub – Cryo EM
    
  • 6jma - XlH3.2 + H4 + H2A + H2B in nucleosome + DOT1 + Ub – Cryo EM
    
  • 7k6p - XlH3.2 (mutant) + H4 + H2A + H2B (mutant) in nucleosome + DOT1 + Ub – Cryo EM
    
  • 6nj9 - XlH3.2 (mutant) + H4 + H2A (mutant) + H2B in nucleosome + DOT1 + Ub – Cryo EM
    
  • 6j99 - XlH3.2 + H4 + H2A + H2B (mutant) in nucleosome + DOT1 + Ub – Cryo EM
    
  • 6nqa - XlH3.2 + H4 + H2A (mutant) + H2B (mutant) in nucleosome + DOT1 + Ub – Cryo EM
    
  • 6o96 - XlH3.2 (mutant) + H4 + H2A (mutant) + H2B (mutant) in nucleosome + DOT1 + Ub – Cryo EM
    
  • 7k6q - XlH3.2 (mutant) + H4 (mutant) + H2A (mutant) + H2B (mutant) in nucleosome + DOT1 + Ub – Cryo EM
    
  • 6ftx - XlH3.1 + H4 + H2A + H2B in nucleosome + CHD1 + Ub – Cryo EM
    
  • 6hts - XlH3.1 + H4 + H2A + H2B in nucleosome + RuvB + CHD1 + actin-related protein – Cryo EM
    
  • 9ge5 - XlHistones in nucleosome + RuvB + INO80 – Cryo EM
    
  • 6ne3 - XlH3.2 (mutant) + H4 + H2A (mutant) + H2B in nucleosome + SNF2H – Cryo EM
    
  • 5mlu - XlH3.2 + H4 + H2A + H2B in nucleosome + PFV GAG peptide
    
  • 5kgf - XlH3.2 + H4 + H2A + H2B in nucleosome + 53BP1 + Ub
    
  • 5hq2 - XlH3.2 + H4 + H2A + H2B in nucleosome + SRM1 + SetD8
    
  • 6ven - XlH3.2 (mutant) + H4 + H2A (mutant) + H2B (mutant) in nucleosome + COMPASS + Ub – Cryo EM
    
  • 6uh5 - XlH3.2 + H4 + H2A + H2B (mutant) in nucleosome + COMPASS + Ub – Cryo EM
    
  • 6z6p - XlH3.2 + H3 + H4 + H2A + H2B in nucleosome + HDA1 – Cryo EM
    
  • 6pa7, 8uw1 - XlH3.2 + H4 + H2A + H2B in nucleosome + DNMT3 – Cryo EM
    
  • 8t9g, 8tas, 8tb9, 8vmj - XlHistones in nucleosome + PRC2 – Cryo EM
    
  • 6wkr - XlH3.2 + H3 + H4 + H2A + H2B in nucleosome + PRC2 + JARID2 – Cryo EM
    
  • 6lth - XlH3.2 + H3 + H4 + H2A + H2B in nucleosome + BAF complex – Cryo EM
    
  • 6zhx, 6zhy, 7enn, 8b0a - XlH3.2 + H3 + H4 + H2A + H2B in nucleosome + ALC1 – Cryo EM
    
  • 6tem - XlH4 + H2A + H2B in nucleosome + CenP-A – Cryo EM
    
  • 7u4d - XlH4 + H2A + H2B in nucleosome + CenP-A + Cenp-N – Cryo EM
    
  • 4zux – XlH3.2 + H4 + H2A + H2B in nucleosome + Saga-associated factors + polyUb + UBP8
    
  • 3mvd - XlH3.2 + H4 + H2A + H2B in nucleosome + regulator of chromosome condensation
    
  • 3tu4 - XlH3.2 + H4 + H2A + H2B in nucleosome + regulatory protein SIR3
    
  • 4r8p - XlH3.2 + H4 + H2A + H2B in nucleosome + polycomb complex protein + ring finger protein
    
  • 6k1p, 6jyl, 6iro, 9jnp, 9jnt, 9jnu, 9jnv, 9jnw, 9jnx, 9jnz, 9jo2, 9jo5, 9liu, 9lj2 – XlHistones in nucleosome + iswI – Cryo EM
    
  • 9dgg – XlHistones in nucleosome + bmi-1 + ring2 – Cryo EM
    
  • 9dg3 – XlHistones in nucleosome + ring1 + Ub – Cryo EM
    
  • 9dby – XlHistones in nucleosome + bmi-1 + ring1 + ring2 + Ub – Cryo EM
    
  • 8hxz, 8ihm – XlHistones + EAF3 - Cryo EM
    
  • 8hxy, 8iht, 8jho, 8kd4, 8kd5, 8kd6, 8kd7, 8tof – XlHistones + Rpd3 + SIN3 + EAF3 + RCO1 - Cryo EM
    
  • 6kw3, 6kw4, 6kw5, 6tda – XlH3 + H4 + H2A + H2B in nucleosome + RSC complex – Cryo EM
    
  • 6rny - hH3.3 + H4 + H2A + H2B in nucleosome + integrase – Cryo EM
    
  • 6t9l – XlH3 + H4 + H2A + H2B in nucleosome + SAGA complex – Cryo EM
    
  • 8g86, 8g88, 8g8b - XlH3 + H4 + H2A + H2B in nucleosome + OCT4 – Cryo EM
    
  • 7vvu - XlH3 + H4 + H2A + H2B in nucleosome + EAF6 +YNG2 + ESA1 – Cryo EM
    
  • 7vvz - XlH3 + H4 + H2A + H2B in nucleosome + EAF6 +YNG2 + ESA1 + NUA4 – Cryo EM
    
  • 7xfh, 7xfm - XlH3.2 + H4 + H2A + H2B in nucleosome + ADPG – Cryo EM
    
  • 8gpn - XlH3.2 + H4 + H2A + H2B in nucleosome + menin – Cryo EM
    
  • 7pgw - XlH3.2 + H4 + H2A + H2B in nucleosome + PC4 and SFRS1-interacting protein – Cryo EM
    
  • 7ssa - XlH3.2 + H4 + H2A + H2B in nucleosome + Cbf1 – Cryo EM
    
  • 8cww – XlHistones in nucleosome + HOP1 – Cryo EM
    
  • 9jao – XlHistones in nucleosome + hHEL1 – Cryo EM
    
  • 9qaj – XlHistones in nucleosome + BCL7A – Cryo EM
    
  • 8g6q, 8v25, 8v26 – XlHistones in nucleosome + Ub – Cryo EM
    
  • 8ett, 8etv, 8eue, 8euj, 9c9g, 9c9t – XlHhistones in nucleosome INO80 complex – Cryo EM
    
  • 9cg9 – XlHistones in nucleosome + HMG-1 - Cryo EM
    
• Yeast nucleosome core + DNA complex with protein
  • 6j51, 6j50, 6j4z, 6j4y, 6j4x, 6j4w, 6inq, 6i84, 6a5u, 6a5t, 6a5r, 6a5p, 6a5o, 6a5l, 7zs9, 7zsa, 7zsb, 8ceo – yH3.3 + H4 + H2A + H2B in nucleosome + RNA polymerase + RNA – Cryo EM
    
  • 6ir9 – yH3.3 + H4 + H2A + H2B in nucleosome + RNA polymerase + SPT4 + SPT5 + RNA – Cryo EM
    
  • 6gen, 6gej – yH3 + H4 + H2A + H2B in nucleosome + SWI1 + actin-like protein + vacuolar protein sorting-associated + RNA – Cryo EM
    
  • 1id3 - yH3 + H4 + H2A + H2B in nucleosome
    
  • 4jjn, 4kud - yH3 + H4 + H2A.2 + H2B.2 in nucleosome + SIR3
    
  • 8qku, 8qkv, 8qyv, 9b1e, 9fbw - yHistones in nucleosome + SWR1 – Cryo EM
    
  • 8xgc - yHistones in nucleosome + replisome – Cryo EM
    
  • 5zba, 5zbb - yH3 + H4 + RTT109 + ASF1
    
  • 8ghm, 8ghn - yH3 + H4 + HIR1+ histone chaperon + histone transcription regulator + histone promoter conrol protein – Cryo EM
    
  • 5z3v, 5z3u, 5z3o, 5z3l - yH3.2 + H4 + H2A + H2B in nucleosome + SNF2 – Cryo EM
    
  • 7k7g - yH3 + H4 + H2A + H2B in nucleosome + CenP-3 – Cryo EM
    
  • 7z0o - yH3 + H4 + H2A + H2B in nucleosome + RNA polymerase transcription initiation factors – Cryo EM
    
  • 7e9c, 7e9f - yH3 + H4 + H2A + H2B in nucleosome + origin recognition complex subunit 1 – Cryo EM
    
  • 6qld, 8ow0 - yH3 + H4 + H2A + H2B in kinetochore + CenP-A – Cryo EM
    
  • 7on1 - yH4 + H2A + H2B in nucleosome + CenP-A + CenP-C – Cryo EM
    
  • 7k78 - yH4 + H2A + H2B in nucleosome + Sce4 + ScfV – Cryo EM
    
  • 8x2x, 8x2y, 8x2z, 8x30, 8x31, 8x32 - yHistones in nucleosome + EAF6 + chromatin modification-related protein + histone acetyltransferase + glutathione transferase – Cryo EM
    
  • 8w9c, 8w9d, 8w9e, 8w9f – yH3 + Rpd3 + SIN3 + EAF3 + RCO1 - Cryo EM
    
• Other nucleosome core + DNA
  • 1u35 - mH3.1 + H2A + H2B + His2A macro domain in nucleosome – mouse
    
  • 5xm1, 5xm0 – mH3 + H4 + H2A + H2B in nucleosome
    
  • 7dbh, 7vbm – mH3mm18 + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 1f66 - XlH3 + mH4 + hH2A + XlH2B in nucleosome
    
  • 5b1m, 5b1l - mHistones in nucleosome
    
  • 8pkj - mHistones in nucleosome – Cryo EM
    
  • 8pki - mHistones in nucleosome + LRH1 – Cryo EM
    
  • 2aro, 1tzy, 1hq3, 2hio, 1hio - cH3 + H4 + H2A + H2B
  • 1eqz - cH3 + H4 + H2A + H2B in nucleosome
    
  • 2pyo, 3nqb, 2nqb - DmH3 + H4 + H2A + H2B in nucleosome
    
  • 6pwe, 9mu4, 9mu5 - DmH3 + H4 + H2A + H2B in nucleosome – Cryo EM
    
  • 6pwf - DmH3 + H4 + H2A + H2B in nucleosome + chromatin remodeling factor – Cryo EM
    
  • 8ux1 - DmHistones in nucleosome + RAN + RCC1 – Cryo EM
    
  • 8pp6, 8pp7 - DmHistones in nucleosome + RYBP + PRC1 – Cryo EM
    
  • 5wcu - DmH3 + H4 + H2A + H2B + H5 in nucleosome
    
  • 4x23 - DmH3 + H4 + H2A + H2B in nucleosome + Cenp-C
    
  • 6dzt - DmH3 + H4 + H2A + H2B in nucleosome + antibody
    
  • 6xws, 6xwt - DmH4 + CenP-A + CAL1
    
  • 8j91, 8j92, 8whb, 9k3z, 9k40, 9k41, 9k42, 9k43 - AtHistones in nucleosome – Cryo EM
    
  • 8j90, 8kcb, 8kcc, 8wh5, 8wh8, 8wh9, 8wha - AtHistones in nucleosome + DDM1 – Cryo EM
    
  • 9mu9 – AtHistones in nucleosome + RNA polymerase complex – Cryo EM
    
  • 8q15 - Histones in nucleosome – rice - Cryo EM
    
  • 6kxv - H3 + H4 + H2A + H2B in nucleosome – Leishmania major
    
  • 7d69 - H3 + H4 + H2A + H2B in nucleosome – Giardia intestinalis – Cryo EM
    
  • 7wlr - H3 + H4 + H2A + H2B in nucleosome – Komagataella pastoris – Cryo EM
    
  • 8com – Histones in nucleosome – Trypanosoma brucei – Cryo EM
    
  • 7lv8, 7lv9 - H3 + H4 + H2A + H2B in nucleosome – Marseillevirus – Cryo EM
    
  • 8ua7 – Histones in nucleosome - medusavirus – Cryo EM
    
  • 7n8n, 9cvt – Histones in nucleosome - melbournevirus – Cryo EM
    
• Chromatosome core + DNA
  • 6la2, 6la8, 6la9, 6lab, 7cow - hH3.1 + H4 + H2A + H2B + H1.0 in chromatosome
    
  • 7dbp - hH3.1 + H4 + H2A + H2B + H1.0 in chromatosome – Cryo EM
    

• • 7k5x, 7k5y, 7k60, 7k61, 7k63 - hH3.1 + H4 + H2A + H2B + H1.0 in chromatosome + scFv – Cryo EM
    
  • 5nl0 - XlH3.2 + H4 + H2A + H2B + H1.0 in chromatosome
    
  • 9ipu - hHistones in chromatosome + RNF168 + Ub-conjugating enzyme + Ub – Cryo EM
    
  • 9dde - hHistones in chromatosome + BMI-1 + Ring1 + Ring2 + Ub – Cryo EM
    
  • 8vg2 – hHistones in chromatosome + GATA4 + FoxA1– Cryo EM

Proteopedia Page Contributors and Editors (what is this?)

Michal Harel, Openfox, Joel L. Sussman