Proteasome 3D structures: Difference between revisions

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**[[8tm4]], [[8tm5]] – h13S – human - Cryo EM<br />
**[[8tm4]], [[8tm5]] – h13S – human - Cryo EM<br />
**[[5dsv]], [h7e55]] – h13S subunit alpha type 3 <br />
**[[5dsv]], [[7e55]] – h13S subunit alpha type 3 <br />
**[[9d4u]] – y13S 7 alpha + 3 beta + UMP1 – yeast - Cryo EM<br />
**[[9d4u]] – y13S 7 alpha + 3 beta + UMP1 – yeast - Cryo EM<br />
**[[7lsx]], [[9rl3]] – y13S 7 alpha + 3 beta + UMP1 + chaperone 1,2 - Cryo EM<br />
**[[7lsx]], [[9rl3]] – y13S 7 alpha + 3 beta + UMP1 + chaperone 1,2 - Cryo EM<br />

Revision as of 09:51, 29 July 2026

3D Structures of Proteasome

Updated on 29-July-2026 {{#tree:id=OrganizedByTopic|openlevels=0|

  • Proteasome 13S core
    • 8tm4, 8tm5 – h13S – human - Cryo EM
    • 5dsv, 7e55 – h13S subunit alpha type 3
    • 9d4u – y13S 7 alpha + 3 beta + UMP1 – yeast - Cryo EM
    • 7lsx, 9rl3 – y13S 7 alpha + 3 beta + UMP1 + chaperone 1,2 - Cryo EM
    • 9rla, 9rlt – y13S 7 alpha + 1 beta + UMP1 + chaperone 1,2 - Cryo EM
    • 9rm0, 9rm1 – y13S 7 alpha + 2 beta + UMP1 + chaperone 1,2 - Cryo EM
    • 9d0t – y13S core 7 alpha + 3 beta + Blm10 – Cryo EM
    • 9d4c – y13S core 7 alpha+ 1 beta + Blm10 – Cryo EM
  • Proteasome Pre-15S core
    • 7ls6 – y15S precursor core - Cryo EM
    • 9rlz – y15S precursor - Cryo EM
  • Proteasome 19S regulatory particle
  • Proteasome 20S catalytic core
  • Proteasome 20S core mutated pre1-1
    • 8rvq – y20S pre1-1 core – Cryo EM
    • 8rvl, 8rvo, 9rvp – y20S precursor pre1-1 core – Cryo EM
    • 9gbk – y20S pre1-1 core + activator – Cryo EM
    • 8t0m – y20S pre1-1, pre4-1 core – Cryo EM
  • Immunoproteasome 20S
  • Preholoproteasome
    • 8qys, 8tm3, 8yix, 8yiy – hPHP-1 – Cryo EM
    • 8yiz – hPHP-2 – Cryo EM
    • 8t08 – yPHP pre1-1, pre4-1 core – Cryo EM
    • 8u6y – yPHP (mutant) – Cryo EM
  • Proteasome 20S+activator protein
  • Proteasome 20S complex
  • Proteasome partial 20S particle
    • 2kqz, 2kr0 – hPRTS ubiquitin receptor domain
    • 2ku1, 2ku2 – Taa-PRTS chains A-G
    • 3fp9 – MtATPase chains A-L
    • 7pxa - MtPRTS α+β subunits + ATPase – Cryo EM
    • 7pxc, 7pxd - MtPRTS α+β subunits + ATPase + Pup – Cryo EM
    • 2jay – MtPRTS β subunit
    • 1j2p – AfPRTS α subunit
    • 5nyw – PRTS β subunit – Yersinia bercovieri
  • Proteasome 26S particle regulatory non-ATPase subunit 1
    • 2n3t - yPRTS regulatory non-ATPase subunit 1 – NMR
    • 2n3u, 2n3v, 2n3w - yPRTS regulatory non-ATPase subunit 1 + L20 – NMR
    • 2nbw - yPRTS regulatory non-ATPase subunit 1 + Rad23 UBL domain – NMR
  • Proteasome 26S particle regulatory non-ATPase subunits 1,2
    • 8usd - hPRTS regulatory non-ATPase subunits 1,2 + Nub1 + regulatory subunits 4,7 – Cryo EM
  • Proteasome 26S particle regulatory non-ATPase subunits 1,13
    • 5v1z, 6oi4 - hPRTS regulatory non-ATPase subunits 1,13 + ubiquitin
    • 6uyi, 6uyj - hPRTS regulatory non-ATPase subunits 1,13 + ubiquitin – NMR
    • 6co4 - hPRTS regulatory non-ATPase subunit 1 peptide + regulatory non-ATPase subunit 13 - NMR
  • Proteasome 26S particle regulatory non-ATPase subunit 2 or Rpn1
    • 7uih - hPRTS regulatory non-ATPase subunit 2 + antibody – Cryo EM
    • 7ujd - hPRTS regulatory non-ATPase subunit 2 + antibody + peptide – Cryo EM


  • Proteasome 26S particle regulatory non-ATPase subunit 4 ofr Rpn10
    • 1p9c – hPRTS regulatory non-ATPase subunit 4 C-terminal
    • 1p9d - hPRTS regulatory non-ATPase subunit 4 C-terminal + HHR23A ubiquitin-like domain
    • 1uel - hPRTS regulatory non-ATPase subunit 4 C-terminal + HHR23A ubiquitin-like domain - NMR
    • 1yx4 - hPRTS regulatory non-ATPase subunit 4 – NMR
    • 1yx5, 1yx6, 2kde, 2kdf, 6mun - hPRTS regulatory non-ATPase subunit 4+ubiquitin – NMR
    • 6u19 - hPRTS regulatory non-ATPase subunit 4 RAZUL domain + ubiquitin protein ligase E3A peptide – NMR
    • 8j4a, 8jtk, 8jtl, 8pfd - PRTS regulatory non-ATPase subunit 4 + SAP05 – Arabidosis thaliana
    • 2x5n - fyPRTS regulatory non-ATPase subunit 4 VWA domain – fission yeast
    • 5ln1 - yPRTS regulatory non-ATPase subunit 4 + polyubiquitin
  • Proteasome 26S particle regulatory non-ATPase subunit 5 or Rpn5
    • 5zmr - yPRTS regulatory non-ATPase subunit 5 N terminal – NMR
  • Proteasome 26S particle regulatory non-ATPase subunit 7 or Rpn9
    • 2o95, 2o96 - hPRTS regulatory non-ATPase subunit 7
    • 3vlf - yPRTS regulatory non-ATPase subunit 7 + DNA mismatch repair protein
    • 2mr3 - yPRTS regulatory non-ATPase subunit 7 – NMR
    • 2mqw - yPRTS regulatory non-ATPase subunit 7 N terminal – NMR
    • 2mri - yPRTS regulatory non-ATPase subunit 7 C terminal – NMR
  • Proteasome 26S particle regulatory non-ATPase subunit 8
    • 3kw6 - hPRTS regulatory non-ATPase subunit 8
    • 2krk - hPRTS regulatory non-ATPase subunit 8 – NMR
  • Proteasome 26S particle regulatory non-ATPase subunits 8,11
    • 4o8x, 4o8y, 4owp, 5w83 - yPRTS regulatory non-ATPase subunits 8,11
    • 4ocl, 4ocm, 4ocn - yPRTS regulatory non-ATPase subunits 8,11 + nanobody
    • 5u4p - yPRTS regulatory non-ATPase subunits 8,11 + S31
  • Proteasome 26S particle regulatory non-ATPase subunit 11
    • 3txm, 3txn – PRTS regulatory non-ATPase subunit 11 – Drosophila melanogaster
    • 6fju, 6fjv, 6fno - CcPRTS regulatory non-ATPase subunit 11 homolog – Candidatus caldiarchaeum
    • 6fnn - CcPRTS regulatory non-ATPase subunit 11 homolog + ubiquitin
  • Proteasome 26S particle regulatory non-ATPase subunit 12
    • 4b0z - fyPRTS regulatory non-ATPase subunit 12
  • Proteasome 26S particle regulatory non-ATPase subunit 13 or ADRM1
    • 4uel - hPRTS regulatory non-ATPase subunit 13 DEUBAD domain 266-388 + UCH-L5 + ubiquitin
    • 2z4d - yPRTS regulatory non-ATPase subunit 13
    • 2z59 - mPRTS regulatory non-ATPase subunit 13+ubiquitin
  • Proteasome 26S particle regulatory non-ATPase subunit 14
    • 3vl1 - yPRTS regulatory non-ATPase subunit 14 (mutant)


  • Proteasome 26S particle Dss1 subunit
    • 3t5x - hPRTS subunit Dss1 + PCI domain-containing protein 2
    • 8yjb - hPRTS subunit Dss1 + INTAC complex – Cryo EM
    • 8yjb, 9vd9 - hPRTS subunit Dss1 + INTAC-PEC complex – Cryo EM
    • 5l4k - hPRTS subunit Dss1 + Lid complex – Cryo EM
  • Proteasome 26S particle SEM1 subunit
    • 28wy, 28xa, 28xb - hPRTS subunit SEM1 + SAC3D1 + PCID2 – Cryo EM
    • 8r7j, 8r7k, 9upc - hPRTS subunit SEM1 in TREX-2 complex – Cryo EM
    • 9dlp, 9t6n, 9opb - hPRTS subunit SEM1 + spliceosome RNA helicase + Germinal-center associated nuclear protein + PCI-domain containing protein – Cryo EM
    • 9dlr, 9dlv, 9t6l - hPRTS subunit SEM1 + spliceosome RNA helicase + leukocyte receptor cluster + PCI-domain containing protein – Cryo EM
    • 9rv1 - hPRTS subunit SEM1 + spliceosome RNA helicase + + PCI-domain containing protein + SAC3 domain containing protein – Cryo EM
    • 3jck - hPRTS subunit SEM1 + Lid complex – Cryo EM
    • 3t5v, 4trq, 5g5p, 5l3t - yPRTS subunit SEM1 + nuclear mRNA export proteins SAC3 and THP1<br /
    • 5ubp - yPRTS subunit SEM1 + nuclear mRNA export protein THP1 + leucine permease transcriptional regulator
    • 7ewf, 7ewm - yPRTS subunit SEM1 + CSN12 + THP3
    • 8u8c - yPRTS subunit SEM1 + SAC3 + THP1 + SUB2
    • 8u8d, 8u8e - yPRTS subunit SEM1 + RNA helicase + nuclear mRNA export factor + THP1 – Cryo EM


For hPRTS regulatory non-ATPase subunit 10 or Gankyrin see Ankyrin

  • Proteasome 26S particle

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Michal Harel

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