ATPase 3D structures

3D Structures of ATPase

ATPase 3D structures

Updated on 29-March-2026

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• ATPase hydrolysing ATP
• F1F0 ATPase α subunit
  • 2jmx - bF1-ATPase+subunit O - bovine
    
  • 6foc – MsF-ATPase – Mycobacterium smegmatis
    
• F1F0 ATPase ε subunit
  • 2rq6, 2rq7 - TeF1-ATPase – Thermosynechococcus elongatus
    
  • 8hgx – AbATPS – Acinetobacter baumannii – NMR
    
• F1F0 ATPase zeta subunit
  • 7vkv - F1-ATPase – Sinorhizobium meliloti - NMR
    
• F1F0 ATPase α, β subunits
  • 1sky – BaF1-ATPase – Bacillus
    
  • 5cdf – PdATPase - Paracoccus denitrificans
    
• F1F0 ATPase α, δ subunits
  • 2a7u – EcF1-ATPase N termini – Escherichia coli - NMR
    
• F1F0 ATPase γ, ε subunits
  • 5zwl – TeF1-ATPase
    
• F1F0 ATPase α, β, γ subunits
  • 1ohh, 2jiz, 4tt3, 4tsf, 4z1m – bF1-ATPase + IF1 inhibitor
    
  • 1e1q, 2w6e, 2w6f, 2w6g, 2w6h, 2w6i, 2w6j - bF1-ATPase
    
  • 1e1r - bF1-ATPase +Mg+ADP+AlF4
    
  • 1w0j, 1w0k - bF1-ATPase +BeF3
    
  • 1nbm - bF1-ATPase +nitrobenzofurazan
    
  • 1mab, 2f43 - rF1-ATPase – rat
    
  • 8hh1, 8hh2, 8hh3, 8hh4, 8hh5, 8hh6, 8hh7, 8hh8, 8hh9, 8hha, 8hhb, 8hhc - BaF1-ATPase – Cryo EM
    
• ”F1F0 ATPase α,β,γ,ε subunits”
  • 1jnv - EcF1-ATPase
    
  • 2qe7, 4xd7 – BaF1-ATPase
    
  • 5ik2, 5hkk – F1-ATPase – Cakdalkalibacillus thermarum
    
  • 6foc – MsF-ATPase
    
  • 6q45 – FnF1-ATPase – Fusobacterium nucleatum
    
  • 7yry, 8zi0, 8zi1, 8zi2, 8zi3 – AbATPS + ADP + ATP – Cryo EM
    
• F1F0 ATPase α, β, γ, δ, ε subunits
  • 4asu, 2jdi, 2jj1, 2jj2 - bF1-ATPase
    
  • 2v7q, 1e79, 2ck3 - bF1-ATPase +inhibitor
    
  • 4yxw - bF1-ATPase + ANP
    
  • 3oe7, 3oee, 3oeh, 3ofn - yF1-ATPase (mutant) - yeast
    
  • 3fks - yF1-ATPase
    
  • 3zia - yF1-ATPase + inhibitor protein
    
  • 6f5d - TbF1-ATPase + ribonucleoprotein P18 – Trypanosoma brucei
    
• F1F0 ATPase multiple subunits
  • 8h9j, 8h9l, 8h9u - hF1-ATPase – Cryo EM
  • 1bmf - bF1-ATPase
    
  • 3dze – bF1-ATPase subunit S
    
  • 1cow - bF1-ATPase +aurovertin B
    
  • 4f4s – yATPase subunit 9 + oligomycin
    
  • 8ap6, 8ap7, 8aph, 8apb - TbF1-ATPase – Cryo EM
  • 7y5a, 7y5b, 7y5c, 7y5d - MsF1-ATPase – Cryo EM
• V-ATPase
• V-ATPase subunit A
  • 7uzg, 7uzh, 7uzi, 7uzk - rATPase + SidK – Cryo EM
  • 5i1m – yATPase
    
  • 6hh0 – yATPase helix 7 - NMR
    
  • 4o1s - ATPase subunit A – Thermoplasma volcanium
    
  • 7qsu – ATPase – Thermococcus litoralis
    
• V-ATPase subunit B
  • 2c61 - MmATPase subunit B – Methanosarcina mazei
    
• V-ATPase subunit C
  • 1u7l - yATPase subunit C
    
  • 1r5z, 1v9m - TtATPase subunit C – Thermus thermophilus
    
• V-ATPase subunit D+F
  • 4rnd – yATPase subunit D+F
    
• V-ATPase subunit E
  • 2kz9 – yATPase subunit E
    
  • 3v6i – TtATPase subunit E + VAPC
    
• V-ATPase subunit F
  • 4ix9 – yATPase subunit F (mutant)
    
• V-ATPase subunit G
  • 2k88 - yATPase subunit G residues 2-58 - NMR
    
  • 2kwy - yATPase subunit G residues 61-101 - NMR
    
• V-ATPase subunit H
  • 1ho8 – yATPase subunit H
    
• V-ATPase subunit K
  • 2bl2, 2cyd - EhATPase subunit K – Enterococcus hirae
    
• V-ATPase subunit I
  • 3rrk – ATPase subunit I N terminal – Meiothermus ruber
    
  • V-ATPase multi subunits
  • 6wlw – hATPase – human - Cryo EM
    
  • 6wlz, 6wm2, 6wm3, 6wm4 – hATPase + ADP – Cryo EM
    
  • 7u4t, 7unf – hATPase + MTOR – Cryo EM
    
  • 6xbw, 6xby – bATPase – Cryo EM
    
  • 7khr – bATPase + antibiotics – Cryo EM
    
  • 7une – bATPase + MTOR – Cryo EM
    
  • 7u8o, 7u8p, 7u8q, 7u8r – pATPase – Cryo EM
    
  • 9b8q - rATPS subunit A,C,E,G,H – Cryo EM
    
  • 9b8o, 9b8p – rATPS + SidK + synaptophysin – Cryo EM
    
  • 6vq6, 6vq7, 6vq8 - rATPS subunit A,B,C,D,E,E2, F,G,K, S1 + effector + renin receptor – Cryo EM
    
  • 6vq9, 6vqa, 6vqb - rATPS subunit A,B,D,E,E2,G,H + effector – Cryo EM
    
  • 6vqc, 6vqg, 6vqh - rATPS subunit A,B,D,E2, F, S1 + renin receptor + RNase K – Cryo EM
    
  • 6vqi, 6vqj, 6vqk - rATPS subunit A,C1,E1,G – Cryo EM
    
  • 9brs, 9bru, 9bry, 9brz – mATPase - Cryo EM
    
  • 7uw9, 7uwa, 7uwb, 7uwc, 7uwd – ATPase - lemon – Cryo EM
    
  • 7fda, 7fdb, 7fdc, 7fde, 7tmm, 7tmo, 7tmp, 7tmq, 7tmr, 7tms, 7tmt – yATPase – Cryo EM
    
  • 7tao, 7tap – yATPase + antibiotics – Cryo EM
    
  • 4dl0, 4efa - yATPase subunits C,G,E
    
  • 3w3a - yATPase subunit A,B,D,F
    
  • 5tj5 - yATPase subunit A,C,C’,C’’,D,E,F
    
  • 5voz, 5voy, 5vox - yATPase subunit A,C,C’,C’’,D,E,F,H – Cryo EM
    
  • 6c6l - yATPase subunit A,C,C’,C’’,D,E,F + V0 assembly protein 1
    
  • 6o7t, 6m0r, 6m0s - yATPase subunit A,C,C’,C’’,E + V0 assembly protein 1 – Cryo EM
    
  • 6o7u - yATPase subunit C,C’,C’’,D,E + V0 assembly protein 1 – Cryo EM
    
  • 6o7v, 6o7w, 6o7x - yATPase subunit A,B,C,C’,C’’,D,E,F,G,H + V0 assembly protein 1 – Cryo EM
    
  • 6pe4, 6pe5 - yATPase subunit A,C,C’,C’’,D,E + V0 assembly protein 1 + cation transporter – Cryo EM
    
  • 5bw9 - yATPase subunit A,B,D,E,G,H – Cryo EM
    
  • 5d80 - yATPase subunit A,B,D,E,F,G,H
    
  • 8eas - yATPase + VMA22 + VMA12P – Cryo EM
    
  • 3a5c, 3a5d - TtATPase subunit A,B,D,F
    
  • 6ly8 - TtATPase subunit A,B,D,F + ADP - Cryo-EM
    
  • 6ly9 - TtATPase subunit C,E,I,K - Cryo-EM
    
  • 5tsj - TtATPase subunit A,B,C,D,E,F,G + antibody - Cryo-EM
    
  • 5gas, 5gar, 7vai, 7vaj, 7vak, 7val, 7vam, 7van, 7vao, 7vap, 7vaq, 7var, 7vas, 7vat, 7vau, 7vav, 7vaw, 7vax, 7vay, 7vb0 - TtATPase subunit A,B,C,D,E,F,G,I - Cryo-EM
    
  • 5y60. 5y5z, 5y5y, 5y5x, 6qum, 6r0w, 6r0y, 6r0z, 6r10 - TtATPase subunit A,B,C,D,E,F,I,K - Cryo-EM
    
  • 3j0j – TtATPase - Cryo EM
    
  • 3gqb - TtATPase subunit A (mutant),B (mutant)
    
  • 3k5b - TtATPase subunit E (mutant)+C
    
  • 5zea, 7dqe - EhATPase subunit A,B (mutant)
    
  • 5ze9 - EhATPase subunit A,B (mutant) + AMPPNP
    
  • 5knd, 5knc, 5knb - EhATPase subunit A,B,D,F
    
  • 8wci - EhATPase subunit K,I – Cryo EM
    
• P-ATPase
• Cu+2 transporting ATPase Domains – 1st metal binding (MBD) 1-79; 2nd metal binding 169-240; 3rd metal binding 275-352; 4th metal binding 375-446; 5th metal binding 486-558; 6th metal binding 562-633; actuator 806-924; nucleotide binding (NBD) 1051-1231
  • 7xuk, 7xun, 7xuo – hATPase (mutant) – Cryo EM
    
  • 7xum – hATPase (mutant) + Cu – Cryo EM
    
  • 8ioy – hATPase (mutant) + AMPPNP – Cryo EM
    
  • 2arf, 2kmv, 2kmx, 2koy – hATPase NBD – NMR
    
  • 2kij - hATPase actuator domain – NMR
    
  • 1yjr, 1yjt, 1yju, 1yjv - hATPase (mutant) 6th MBD – NMR
    
  • 1y3j, 1y3k - hATPase 5th MBD – NMR
    
  • 2ew9 - hATPase 5th+6th MBDs – NMR
    
  • 1q8l, 1s6o, 1s6u, 2lqb - hATPase 2nd MBD– NMR
    
  • 1aw0, 2aw0 - hATPase 4th MBD – NMR
    
  • 1kvi, 1kvj - hATPase 1st MBD – NMR
    
  • 5t7l - hATPase 1st MBD + Cu transport protein
    
  • 2k1r - hATPase 1st MBD+ ATOX1 – NMR
    
  • 2rop, 2g9o, 2ga7 - hATPase 3rd MBD – NMR
    
  • 1fvq, 1fvs - hATPase – NMR
    
  • 3cjk – hATPase + Cu transporting protein ATX1
    
  • 3dxs - AtATPase RAN1 MBD (mutant) – Arabidopsis thaliana
    
  • 5lbd – AtATPase PAA1 N-domain
    
  • 5lbk – AtATPase PAA2 N-domain
    
  • 8q76 – riATPase – rice – Cryo EM
    
  • 8q73, 8q75 - riATPase 101-978 – Cryo EM
    
  • 8q74 - riATPase 101-978 + Cu – Cryo EM
    
  • 3voy, 3j08, 3j09 - AfATPase – Archaeoglobus fulgidus – CryoEM
    
  • 3skx, 3sky - AfATPase NBD
    
  • 3a1c – AfATPase NBD + AMPPCP
    
  • 3a1d - AfATPase NBD + ADP-Mg
    
  • 3a1e - AfATPase NBD (mutant) + AMPPCP
    
  • 2rml - BsATPase N-terminal – Bacillus subtilis – NMR
    
  • 2gcf - syATPase N-terminal - Synechocystis – NMR
    
  • 4a4j - syATPase N-terminal
    
  • 4a48 - syATPase
    
  • 2iye - SsATPase catalytic domain – Sulfolobus solfataricus
    
  • 2yj3 - SsATPase catalytic domain (mutant)
    
  • 2yj4, 2yj5, 2yj6 - SsATPase catalytic domain (mutant) + nucleotide
    
  • 3rfu, 4bbj, 4bev, 4byg – LpATPase – Legionella pneumophila
    
  • 8ovl – LpATPase 1st MBD – NMR
    
  • 4f2f - ATPase MBD – Streptococcus pneumonia
    
• Na/K transporting ATPase
  • 7e1z, 7e20, 7e21 - hATPase – Cryo EM
    
  • 8d3u, 8d3v, 8d3w, 8d3x, 8d3y - hATPase + ion transport regulator – Cryo EM
    
  • 3kdp, 3b8e, 7yzr – pATPase - pig
    
  • 3n23 – pATPase + ouabain
    
  • 3wgv, 3wgu – pATPase + oligomycin
    
  • 3b8e - pATPase a+BeF3
    
  • 1q3i - pATPase NBD
    
  • 8ijv - pATPase subunits α,β
    
  • 4ret, 4res, 4hyt, 4hqj, 8jbl, 8jbm - pATPase subunits α,β,γ
    
  • 7d91, 7ddf, 7wyt, 7qtv, 7z04 - pATPase subunits A,B + ion transport regulator
    
  • 7d92, 7d93, 7d94, 7wys, 7ddh, 7ddi, 7ddk, 7ddl - pATPase subunits A,B + ion transport regulator + inhibitor
    
  • 8ijw, 8jmn - pATPase subunit A + K transport ATPase subunit B – Cryo EM
    
  • 8wa5 - pATPase subunit A (mutant) + K transport ATPase subunit B + K – Cryo EM
    
  • 8ijx - pATPase subunit A (mutant) + K transport ATPase subunit B + inibitor – Cryo EM
    
  • 8jbk - pATPase subunits A,B + K transport ATPase subunit C
    
  • 4xe5 - bATPase
    
  • 1mo8, 1mo7 - rATPase α-1
    
  • 8ijl - rATPase A,B – Cryo EM
    
  • 8ijm - rATPase A,B (mutan t) + Na – Cryo EM
    
  • 2hc8 – AfATPase CopA A domain
    
  • 2b8e - AfATPase CopA NBD
    
  • 3a3y - sdATPase+K+ouabain – spiny dogfish
    
  • 2xze - sdATPase
    
  • 2zxe, 5aw9, 5aw8, 5aw7, 5aw6, 5aw5, 5aw4, 5aw3, 5aw2, 5aw1, 5aw0, 5avz, 5avy, 5avx, 5avw, 5avv, 5avu, 5avt, 5avs, 5avr, 5avq – sdATPase + phospholemman-like protein
    
  • 7wyu, 7wyv, 7wyw, 7wyx, 7wyy, 7wyz, 7wz0, 7y45, 7y46, 8jfz- sdATPase + ion transport regulator – Cryo EM
    
  • 8k1l- ATPase – Artemia salina - Cryo EM
    
• Ca+2 transporting ATPase
  • 7yam, 8iwr, 8iws, 8iwt, 8iwu - hATPase – Cryo EM
    
  • 6a69 - hATPase + neuroplastin – Cryo EM
    
  • 8iww - hATPase + ADP – Cryo EM
    
  • 8iwp - hATPase + Ca – Cryo EM
    
  • 7yag, 7yai, 7yaj - hATPase + nanobody + ACP – Cryo EM
    
  • 3fgo - raATPase+CPA+AMPPCP – rabbit
    
  • 3b9r, 1xp5 - raATPase+AlF4
    
  • 1wpg - raATPase+MgF4
    
  • 3w5b - raATPase+Mg
    
  • 4h1w, 3w5a - raATPase+ sarcolipin
    
  • 3w5c, 3w5d, 1iwo - raATPase
    
  • 4bew - raATPase+ phosphate analog
    
  • 2zbe, 3b9b – raATPase+BeF3
    
  • 2zbf - raATPase+BeF3+TG
    
  • 2zbg - raATPase+AlF4+TG
    
  • 2zbd, 3ar8 - raATPase+AlF4+nucleotide+Ca
    
  • 3ar9 - raATPase+BeF3+nucleotide
    
  • 2dqs, 3ar2, 1vfp, 4xou - raATPase+AMPPCP
    
  • 5a3r - raATPase + BeF3 + AMPPCP
    
  • 2c88, 2c8k, 3ar4, 3ar3, 3ar5, 3ar7 - raATPase+nucleotide+TG
    
  • 2ear, 2c8l - raATPase+TG
    
  • 2agv - raATPase+TG+BHQ
    
  • 4ycl - raATPase+CPA
    
  • 2eat - raATPase+CPA+GT
    
  • 3ar6 - raATPase+TNP-ADP+GT
    
  • 2eau - raATPase+CPA+curcumin
    
  • 3ba6 - raATPase phosphoenzyme intermediate
    
  • 3fpb - raATPase+ATP+cyclopiazonic acid
    
  • 3fps, 2oa0 - raATPase+ADP+cyclopiazonic acid
    
  • 2o9j - raATPase+MgF4+cyclopiazonic acid
    
  • 1kju, 3n5k - raATPase E2 state
    
  • 2c9m - raATPase Ca2E1 state
    
  • 1t5s, 3n8g - raATPase Ca2E1 state+AMPPCP
    
  • 4nab - raATPase Ca2E1 state (mutant) +AMPPCP
    
  • 1t5t - raATPase Ca2E1 state+ADP+AlF4
    
  • 1su4 - raATPase +2Ca2
    
  • 2by4, 2yfy, 3nal, 3nam, 3nan, 4uu1, 4uu0, **4j2t, 5a3s - raATPase HNE2 state+thapsigargin derivative
    
  • 5a3q - raATPase + thapsigargin derivative + AMPPCP
    
  • 4ycn, 4ycm - raATPase + macrolide
    
  • 4y3u - raATPase + phospholamban
    
  • 8qmp - AtATPase+BeF3
    
  • 6zhf, 6zhg - LmATPase – Listeria monocytogenes
    
  • 6zhh - LmATPase (mutant) + phosphocholine
    
  • 9gqo - LmATPase + Ca – Cryo EM
    
• “Zn+2 ATPase
  • 2ofg, 2ofh - sATPase N-terminal – NMR
    
  • 4umw, 4umv - ATPase +Mg – Shigella sonni
    
• “Cd+2 ATPase
  • 7qbz, 7qc0 - ATPase – Sulfitobacter
    
• K+ transporting ATPase
  • 2ynr, 4ux2, 4ux1, 5y0b – pATPase – Cryo EM
    
  • 5ylv – pATPase + pyridine derivative
    
  • 5ylu, 7w47, 7w48, 7w49, 7w4a – pATPase + drug
    
  • 6jxh – pATPase (mutant) + K
    
  • 6jxk – pATPase + Rb
    
  • 6jxi, 6jxj – pATPase (mutant) + Rb
    
  • 7et1, 7efl, 7efm, 7efn – pATPase subunit beta + Na/K ATPase subunit alpha (mutant) + cholesterol – Cryo EM
    
  • 7x20, 7x21, 7x22, 7x23, 7x24 – raATPase subunit beta + Na/K ATPase subunit alpha (mutant) – Cryo EM
    
  • 6hra, 6hrb, 7bgy, 7bh1, 7bh2, 7lc3, 7lc6, 7nnl, 7nnp – EcATPase – Cryo EM
    
  • 7zre, 7zrg, 7zrh, 7zri, 7zrj, 7zrk, 7zrm – EcATPase + K – Cryo EM
    
  • 7zrd – EcATPase + K + inhibitor – Cryo EM
    
  • 2a00, 2a29 - EcATPase NBD of KdpB+AMPPNP – NMR
    
  • 1svj, 1u7q - EcATPase NBD of KdpB – NMR
    
• As+ transporting ATPase or GET3 or ATPase asna1 homolog
  • 7ru9, 7ruc - ATPase GET3 (mutant) + golgi to ER traffic protein + ubiquitin-like protein + BAG6 – zebrafish – Cryo EM
    
  • 3h84, 3idq, 3a36, 3a37 - yATPase GET3
    
  • 4pwx, 5bwk - yATPase GET3 + golgi to ER traffic protein + ubiquitin-like protein
    
  • 5bw8 - yATPase GET3 + golgi to ER traffic protein + ubiquitin-like protein + protein
    
  • 4xwo - yATPase GET3 + antibody + Sec22
    
  • 4xvu - yATPase GET3 + antibody + NYV1
    
  • 4xtr - yATPase GET3 + antibody + pep12p
    
  • 3sja, 3sjb, 3sjc, 3sjd, 3zs8, 3zs9, 3b2e, 3vlc - yATPase GET3 + GET1 cystolic domain
    
  • 1ii0, 1f48, 1ii9 - EcATPase
    
  • 1ihu – EcATPase+Mg+ADP+AlF3
    
  • 3ibg - ATPase GET3 – Aspergillus fumigatus
    
  • 8odv - ATPase GET3 (mutant) + GET2 + GET1 – Thermochaetoides thermophila
    
  • 8had - PtATPase GET3 – Phaeodactylum tricornutum
    
  • 8hac - PtATPase GET3 + ADP
    
  • 7spz - GlATPase GET3 – Giardia lamblia
    
  • 7spy - GlATPase GET3 + ATP
    
  • 7sq0 - GlATPase GET3 + ADP
    
• H+ transporting ATPase
  • 7vh5, 7vh6 – yATPase - Cryo EM
    
  • 5ksd - AtATPase C terminal truncated
    
  • 1mhs - NcATPase – Neurospora crassa
    
  • 7op1, 7op3, 7op5, 7op8, 8odu – CtATPase – Chaetomium thermophilum - Cryo EM
    
• Na+ transporting ATPase
  • 2db4, 3aou - EhATPase subunit K
    
  • 3aon - EhATPase subunits D,G
    
  • 3vr2, 8igu, 7coq, 8igv, 8igw - EhATPase subunits A,B
    
  • 3vr4, 3vr5 - EhATPase subunits A,B,D,G
    
  • 3vr3 - EhATPase subunits A,B + AMPPNP
    
  • 3vr6 - EhATPase subunits A,B,D,G + AMPPNP
    
  • 1yce - IlATPase rotor ring
• H/K transporting ATPase
  • 1iwc, 1iwf - pATPase NBD
    
  • 3ixz - pATPase a+AlF4
    
  • 2xzb – pATPase subunits α, β
    
• Mg+2 transporting ATPase
  • 3gwi – EcATPase P-1 NBD
• Arg/ornithine transporting ATPase
  • 1h8e – bATPase+ADP+AlF4+ADP+SO4
    
  • 1h8h -bcATPase+ AMPPNP
    
  • 1efr - bATPase+efrapeptin
    
  • 3cf0, 3cf1, 3cf3 – mATPase NBD+ADP – mouse
    
  • 3cf2 - mATPase NBD+ADP+AMP-PNP
    
  • 1kmh - sATPase alpha subunit+tentoxin
    
  • 3fks – yATPase
    
  • 3ea0 – ATPase ParA, fragment – Chlorobium tepidum
    
  • 2qen – ATPase (mutant) Walker-type – Pyrococcus abyssi
    
  • 2r31, 2p4x – PdATPase ATP12
    
  • 2zd2 – PdATPase (mutant) ATP12
    
  • 1d8s – EcATPase F1
    
  • 1vdz – PhATPase subunit A – Pyrococcus horikoshii
    
  • 3md0 – MtATPase+GDP
    
• Monosaccharide transporting ATPase
  • 5dte – AsATPase – Actinobacillus succinogenes
    
  • 4zjp – AsATPase + ribopyranose
    
• Phospholipid transporting ATPase
  • 7lkp, 7roq, 7tdt, 7w01, 8edw, 8ee6, 8eeb – hATPase – Cryo EM
    
  • 7w02, 8eop – hATPase (mutant) + ATP – Cryo EM
    
  • 6k7g, 6k7h, 6k7i, 6k7j, 6k7k, 6k7l, 6k7m, 6k7n, 7py4, 7vgh, 7vgi, 7vgj, 7vsg, 7vsh, 8ox4, 8ox5, 8ox6, 8ox7, 8ox8, 8ox9, 8oxa, 8oxc – hATPase + cell cycle control protein – Cryo EM
    
  • 6lkn – hATPase (mutant) + cell cycle control protein
    
  • 7drx, 7f7f, 7ky5, 7ky6, 7ky7, 7kya, 7kyc – yATPase + LEM3 – Cryo EM
    
  • 7ky9, 7kyb – yATPase + LEM3 + ADP – Cryo EM
    
  • 7dsh, 7dsi, 7ky8, 7whv, 7whw – yATPase + LEM3 + AMPPCP – Cryo EM
    
  • 6lcp, 6lcr – CtATPase + cell cycle control protein - Cryo EM
    
• Polyamine transporting ATPase
  • 7vpi, 7vpk, 8iel, 8ien, 8ieo – hATPase – Cryo EM
    
  • 7vpj, 8ies – hATPase + ADP – Cryo EM
    
  • 8iek – hATPase + ATP – Cryo EM
    
  • 7vpl, 8iem – hATPase + spermine – Cryo EM
    
• RNA-dependent ATPase
  • 3eaq, 3i31, 3i32 – TtATPase fragment
    
  • 4kbg, 4kbf - TtATPase helicase core domain
    
  • 5mao - TtATPase RRM domain
    
  • 4i69, 4i68 - TtATPase RRM domain (mutant)
    
  • 4i67 - TtATPase RRM domain + RNA
    
  • 3mwj – TtATPase N-terminal (mutant)
    
  • 3mwk, 3nbf – TtATPase N-terminal + 8-oxo-AMP
    
  • 3mwl - TtATPase N-terminal (mutant) + 8-oxoadenine
    
  • 3nej - TtATPase reca-like domain (mutant)
    
• DNA-dependent ATPase
  • 5tnu - ATPase – Sulfolobus tokodaii
    
  • 6p4o, 6p4w - ATPase + endonuclease BAX1 – Sulfurisphaera tokodaii
    
• Proteasome-associated ATPase
  • 7qxu – hATPase in proteasome – Cryo EM
    
  • 8d6v, 8d6w – MtATPase subunits alpha, beta – Cryo EM
    
  • 7pxc – MtATPase subunits alpha, beta + Pup – Cryo EM
    
  • 3fp9, 3m9b, 3m9h – MtATPase intern domain
    
  • 3m91, 3m9d - MtATPase coiled coil domain + protein PUP
    
  • 5kzf – MtATPase residues 98-609
    
  • 5kwa – MtATPase residues 22-529
    
  • 6kww – ATPase subunit HsIU – Staphylococcus aureus
    
  • 5txv – EcATPase subunit HslU
    
  • 6pxk, [[[6pxl|6pxl]], 6pxx – EcATPase subunit HsIU + ADP
    
  • 5ji3, 5ji2 – EcATPase subunits HslU+HslV
    
  • 6pxi – EcATPase subunit HsIU+HsIV + ADP
    
  • 2wfw – ATPase substrate recognition domain – Rhodococcus erythropolis
    
  • 7mhb – ATPase subunit HsIU + ADP – Brucella abortus
    
• Transitional endoplasmic reticulum ATPase residues 1-806
  • 7r7u – hTer-ATPase – Cryo EM
    
  • 5c19 – hTer-ATPase (mutant)
    
  • 8pqx, 8r0e, 8rs9 – hTer-ATPase (mutant) – Cryo EM
    
  • 5ftn, 5ftm, 5ftl, 5ftk, 5ftj – hTer-ATPase + ATPGS
    
  • 7vcs, 7vcv, 7vcx – hTer-ATPase + ATPGS – Cryo EM
    
  • 5c18 – hTer-ATPase (mutant) + ATPGS
    
  • 7rl7, 7rla, 7rlc, 7rlf, 7rlh – hTer-ATPase (mutant) + ATPGS – Cryo EM
    
  • 7rlj – hTer-ATPase (mutant) + ATPGS + inhibitor – Cryo EM
    
  • 7rli – hTer-ATPase + ADP + inhibitor – Cryo EM
    
  • 7wub, 8ooi, 8rsb – hTer-ATPase + ADP – Cryo EM
    
  • 7rl6, 7rl9, 7rlb, 7rld, 7rlg, 8rsc – hTer-ATPase (mutant) + ADP – Cryo EM
    
  • 7vct, 7vcu – hTer-ATPase + ATPGS + ADP – Cryo EM
    
  • 8fcl, 8fcm, 8fcn, 8fco, 8fcp, 8fcq, 8fcr, 8fct – hTer-ATPase + UBXD1 + ADP – Cryo EM
    
  • 7y4w, 7y53, 7y59 – hTer-ATPase + derlin + ADP – Cryo EM
    
  • 8b5r – hTer-ATPase + P37 + SPI – Cryo EM
    
  • 5c1b - hTer-ATPase + UFD1--SHP peptide
    
  • 5ifw, 5ifs - hTer-ATPase + tether containing UBX domain
    
  • 6mck - hTer-ATPase + inhibitor
    
  • 8uv2, 8uvp, 9boq - hTer-ATPase + inhibitor – Cryo EM
    
  • 8uvq - hTer-ATPase (mutant) + inhibitor – Cryo EM
    
  • 8uvo - hTer-ATPase (mutant) + ADP + inhibitor – Cryo EM
    
  • 6hd0 - hTer-ATPase + PUX1 + ADP
    
  • 8hl7 - hTer-ATPase + N-Lys methyltransferase + SAH + ADP
    
  • 8yka - hTer-ATPase + deubiquitinqting protein – Cryo EM
    
  • 7r7s - hTer-ATPase + p47 – Cryo EM
    
  • 7r7t - hTer-ATPase + p47 + ADP – Cryo EM
    
• Transitional endoplasmic reticulum ATPase residues 1-480
  • 3hu1, 3hu2, 3hu3 – hTer-ATPase + ATPGS
    
  • 4ko8, 4kln, 5dyg – hTer-ATPase (mutant) + ATPGS
    
  • 5dyi – hTer-ATPase + ADP
    
  • 4kod – hTer-ATPase (mutant) + ADP
    
  • 5kiy, 5kiw - hTer-ATPase (mutant) + Selenoprotein S
    
  • 8kg2 – hTer-ATPase (mutant) + FAS-associated factor
    
  • 8hrz – hTer-ATPase (mutant) + P47 UBX domain
    
• Transitional endoplasmic reticulum ATPase residues 1-200
  • 3qc8, 3qq8, 3qwz - hTer-ATPase + FAS-associated factor 1
    
  • 3qq7 - hTer-ATPase
    
  • 3tiw - hTer-ATPase + E3 ubiquitin-protein ligase peptide
  • 5glf - hTer-ATPase + derlin-1 peptide
    
  • 5epp - hTer-ATPase N + rhomboid-related protein 4 peptide
    
  • 4kdl, 4kdi - hTer-ATPase + ubiquitin thioesterase OTU1
    
  • 5b6c - hTer-ATPase + ubiquitin fusion degradation protein
    
  • 5x4l - hTer-ATPase + UBX domain-containing protein
    
  • 1s3s - mTer-ATPase ND1 domain + P47 C terminal
    
  • 4rv0 - Ter-ATPase N terminal + nuclear protein localization protein – Drosophila melanogaster
    
• Transitional endoplasmic reticulum ATPase D2 domain residues 462-764
  • 6g2v, 6g2w, 6g2x, 6g2y, 6g2z, 6g30 - hTer-ATPase + ADP
    
• Sarcoplasmic/endoplasmic reticulum ATPase
  • 6jju, 5ztf, 7bt2 – hATPase 2
    
  • 7w7u, 7w7v, 7w7w – hATPase 2 – Cryo EM
    
  • 7w7t – hATPase 2 + Ca – Cryo EM
    
  • 6lle, 6lly, 6ln5, 6ln6, 6ln7, 6ln8, 6ln9 – hATPase 2 + AMPPCP – Cryo EM
    
  • 5mpm, 6hxb – pATPase
    
  • 5xa7, 5xa8, 5xa9, 5xaa, 5xab, 6yso – raATPase
    
  • 5zmw – raATPase (mutant)
    
  • 5ncq – raATPase + ATP derivative + carbazole derivative
    
  • 6hef, 6rb2 – raATPase + AMPPCP
    
  • 8owa, 8owl– raATPase + thapsigargin
    
  • 5zmv – raATPase (mutant) + thapsigargin
    
  • 6yaa – raATPase + ATP derivative + inhibitor
    
• LAO/AO transport system ATPase
  • 3nxs – MsATPase + GDP
    
  • 6cum – MsATPase
    
• DNA double-strand repair ATPase (Rad50)
  • 3aux – MjRad50 – Methanocaldococcus jannaschii
    
  • 3auy - MjRad50 + ADP
    
  • 3av0 - MjRad50 + ATP
    
  • 5f3w - MjRad50 + MRE11 + ATPGS
    
  • 5dny - MjRad50 + MRE11 + DNA
    
  • 3qg5 – TmRad50 NBD + MRE11
    
  • 3qkr, 3qks, 3qf7 - PfRad50 NBD + MRE11 – Pyrococcus furiosus
    
  • 4nck, 4nci, 4nch - PfRad50 NBD (mutant)
    
  • 4ncj - PfRad50 NBD (mutant) + BeF3 + ADP
    
  • 3qkt - PfRad50 NBD + AMPPNP
    
  • 3qku - PfRad50 NBD + AMPPNP + MRE11
    
• MipZ ATPase
  • 2xit, 2xj4 – CcMipZ – Caulobacter crescentus
    
  • 2xj9 – CcMipZ (mutant)
    
• ATPase Rava
  • 5fl2, 4upf – EcATPase LARA domain + lysine decarboxylase
    
  • 4upb, 6q7l, 6q7m – EcATPase + lysine decarboxylase – Cryo EM
    
  • 3nbx – EcATPase Rava + ADP
    
  • 6sza, 6szb – EcATPase Rava + ADP – Cryo EM
    
• FlaI ATPase
  • 4ihq, 4ii7 – ATPase (mutant) – Sulfolobus acidocaldarius
    
• Vescicle-fusing ATPase
  • 6mdm, 6mdn – haATPase + SNAP-25 + syntaxin-1A + VAMP-2 + NSF attachment protein – hamster – Cryo EM
    
  • 6mdo, 6mdp – haATPase + SNAP-25 – Cryo EM
    
• ATPase Invc
  • 6sdx – ATPase – Salmonella enterica
    
• ATPase in ribosome
  • 6tmf – ATPase – Thermococcus celer
    
• P5A-type ATPase
  • 6xmp, 6xmq, 6xms, 6xmt, 6xmu – yATPase – Cryo EM
    
• ATPase SpaL/MxiB
  • 6n6l, 6n6m, 6n70, 6n71, 6n72, 6n73, 6n75, 6n76 – ATPase (mutant) – Shigella flexneri
    
• Plasma membrane ATPase
  • 7nxf, 7ny1 – NcATPase – Cryo EM
    
• Peroxisomal ATPase
  • 8u0x – yPex6 N1 domain 1-184
    
  • 8u0v – yPex1 + Pex6 – Cryo EM
    
  • 8c0v, 8c0w – yPex1 + Pex6 (mutant) + peptide + ADP + ATP – Cryo EM
    
• Archaeal ATPase
  • 8fnw, 8fnt – EcATPase – Cryo EM
    
  • 8hr7, 8hr8, 8hr9 – EcATPase + ATP – Cryo EM
    
  • 8hra, 8hrb – EcATPase + ATP + RNA – Cryo EM
    
• Ino80 ATPase or chromatin-remodeling ATPase
  • 6hts, 7vdv, 7y8r, 7zi4, [[[8ooa|8ooa]], [8x15]], 8x19, 8x1c – hIno80 in nucleosome – Cryo EM
    
  • 8ett, 8etv, 8eu2, 8eue, 8euj – fIno80 in nucleosome – frog - Cryo EM
    
  • 6gej, 6gen, 7x3t, 8a5a, 8a5o, 8ets, 8etu, 8etw, 8eu9, 8euf, 8qku, 8qkv, 8qyv, 8qz0, 9b1d, 9b1e, 9fbw – yIno80 in nucleosome – Cryo EM
    
  • 5nbn – yIno80 in nucleosome
    
  • 8a5d, 8a5p, 8a5q, 8atf – CtIno80 in nucleosome – Cryo EM
    
  • 6fml, 8atf, 8av6, 8oo7, 8oo9, 8ooc, 8oof, 8oop, 8oor, 8oos, 8oot – TtIno80 + DNA – Cryo EM
    
• ATPase synthesizing ATP or ATP synthase
• V-ATPS subunit A
  • 3i4l, 7qss, 7qst - PhATPS subunit A
    
  • 3mfy, 3m4y, 3nd8, 3nd9, 3p20, 3qg1, 3qia, 3qjy, 3sdz, 3se0, 5x09 - PhATPS subunit A (mutant)
  • 3i72, 3i73, 3ikj - TtATPS subunit A (mutant)
    
  • 5bn5 – NeATPS + NEQ263 – Nanoarchaeum equitans
    
  • 5bn4 – NeATPS + NEQ263 + ANP
    
  • 5bn3 – NeATPS + NEQ263 + ADP
    
  • 5bo5 – NeATPS subunit NeqB
    
• V-ATPS subunit B
  • 2rkw, 3b2q, 3ssa, 3tgw, 3tiv - MmATPS subunit B (mutant)
    
  • 3dsr - MmATPS subunit B+ADP
    
  • 3eiu - MmATPS subunit B+ATP
    
• V-ATPS subunit C
  • 3v3c – ATPS subunit C – pea
    
  • 4mjn - F1-ATPS - wheat
    
  • 2w5j, 6tqj – sF1-ATPS - spinach
    
  • 3u2f, 3u2y, 3u32, 3ud0 - yATPS subunit C
    
  • 1ijp, 1l6t – EcF1-ATPS subunit C (mutant) - NMR
    
  • 4utq – EcF1-ATPS membrane domain + insertase YIDC – Cryo EM
    
  • 3lg8 - MjATPS subunit C C-terminal
    
  • 2wie, 2xqs, 2xqt, 2xqu - ATPS subunit C – Arthrospira platensis
    
  • 3zk1, 3zk2 - ATPS subunit C
    
  • 7wem – BaF1-ATPS subunit C - NMR
    
  • 4bem - F1F0-ATPS – Acetobacterium woodii
    
• V-ATPS subunit E
  • 2dm9, 2dma - PhATPS subunit E
    
  • 4dt0 – PhATPS subunit E
    
  • 2kk7 - MjATPS subunit E N-terminal – NMR
    
• V-ATPS subunit F
  • 2ov6 - MmATPS subunit F - NMR
    
  • 2d00 - TtATPS subunit F
    
  • 2qai - PfATPS subunit F
    
  • 2i4r - AfATPS subunit F
    
• F1F0 ATPS α subunit
  • 2r9v - F1-ATPS – Thermotoga maritima
    
• F1F0 ATPS β subunit
  • 1pyv - F1-ATPS – NMR – tobacco
    
  • 1b9u – EcF1-ATPS membrane domain – NMR
    
  • 1l2p – EcF1-ATPS dimerization domain
    
  • 2khk - EcF1-ATPS fragment – NMR
    
  • 4q4l - F1-ATPS – Burkholderia thailandensis
    
• F1F0 ATPS γ subunit
  • 1wu0 – BaATPS – NMR
    
  • 1a91, 1c0v, 1aty, 1c99 – EcF1-ATPS – NMR
    
  • 2wgm – ItF0-ATPS – Ilyobacter tartaricus
    
  • 4v1g – MpF0F1-ATPS – Mycobacterium phlei
    
  • 4v1f, 4v1h – MpF0-ATPS + drug
    
  • 2x2v, 4cbk, 4cbj – BpF0-ATPS – Bacillus pseudofirmus
    
  • 3zo6 – BpF0-ATPS (mutant)
    
• F1F0 ATPS δ subunit
  • 1abv – EcF1-ATPS N terminal – NMR
    
• F1F0 ATPS ε subunit
  • 1aqt – EcF1-ATPS (mutant)
    
  • 1bsh, 1bsn – EcF1-ATPS – NMR
    
  • 2e5t, 2e5u – BaF1-ATPS C terminal – NMR
    
  • 2e5y – BaF1-ATPS
    
  • 6pi4 – MsF1-ATPS
    
  • 7vil, 7xkz – F1-ATPS – Mycobacterium abscessus - NMR
    
  • 5yio – MtF1-ATPS+GDP – Mycobacterium tuberculosis - NMR
    
• F1F0 ATPS α,b subunits
  • 8spv – BaF1-ATPS – Cryo EM
    
  • 8spx – BaF1-ATPS + ATP – Cryo EM
    
  • 8spw – BaF1-ATPS + ATP + ADP – Cryo EM
    
• F1F0 ATPS α,γ subunits
  • 1c17 – EcF1-ATPS – NMR
    
  • 8j58, 8jr1 – MtF1-ATPS – Cryo EM
    
  • 8j57 – MtF1-ATPS + drug – Cryo EM
    
• F1F0 ATPS β,δ subunits
  • 8h9g, 8h9n – hF1-ATPS + ATPS coupling factor – Cryo EM
    
  • 2cly – bF1-ATPS + ATPS coupling factor
    
• F1F0 ATPS γ,ε subunits
  • 1fs0 – EcF1-ATPS
    
• F1F0 ATPS α,β,γ,δ subunits
  • 1qo1- yF1-ATPS + protein 9
    
• F1F0 ATPS α,β,γ subunits
  • 9avj - BaF1-ATPS + ANP – Cryo EM
    
  • 8u1h - BaF1-ATPS (mutant) + ANP + ADP – Cryo EM
    
• ATPS multi subunits
  • 8h9e, 8h9f, 8h9i, 8h9k, 8h9m, 8h9p, 8h9q, 8h9r, 8h9s, 8h9t, 8h9v – hATPS – Cryo EM
    
  • 8khf, 8ki3 – hATPS + drug – Cryo EM
    
  • 5fil, 5fik, 5fij, 5ari, 5arh, 5are, 5ara, 6xbw, 6xby, 6zbb, 6zg7, 6zg8, 6zik, 6ziq, 6zit, 6ziu, 6zpo, 6zqm, 6zqn, 7ajb, 7ajc, 7ajd, 7aje, 7ajf, 7ajg, 7ajh, 7aji, 7ajj – bATPS – Cryo EM
    
  • 4asu, 2jdi, 2jj1, 2jj2 - bF1-ATPS
    
  • 2v7q, 1e79, 2ck3, 2zia - bF1-ATPS+inhibitor
    
  • 6yy0, 6z1r, 6z1u - bF1-ATPS+ inhibitor + coupling factor + ADP + ATP – Cryo EM
    
  • 2xnd, 2xok - bF1-ATPS+ATPS lipid-binding protein
    
  • 6j54, 6j5a, 6j5i, 6j5j, 6j5k, 6zna, 6zmr – pATPS - Cryo EM
    
  • 6j5k – pATPS + ADP + ATP - Cryo EM
    
  • 6za9 – oATPS – ovine - Cryo EM
    
  • 6tt7 – oATPS + ADP - Cryo EM
    
  • 6vm1, 6vm4, 6vmg – sATPS - Cryo EM
    
  • 6fkf, 6fkh, 6fki, 6vmb, 6vmd, 6vof, 6vog, 6voh, 6voi, 6voj, 6vok, 6vol, 6vom, 6von, 6voo – sATPS + ADP + ATP - Cryo EM
    
  • 4b2q, 6b2z, 6b8h, 6cp6, 6cp7,6wtd, 7tjy, 7tjz, 7tk0, 7tk1, 7tk2, 7tk3, 7tk4, 7tk5, 7tk6, 7tk7, 7tk8, 7tk9, 7tka, 7tkb, 7tkc, 7tkd, 7tke, 7tkf, 7tkg, 7tkh, 7tki, 7tkj, 7tkk, 7tkl, 7tkm, 7tkn, 7tko, 7tkp, 7tkq, 7tkr, 7tks, 8eat, 8eau, 8f29, 8f39, 8fl8 – yATPS – Cryo EM
    
  • 3fks, 3zry, 2hld - yF1-ATPS
    
  • 3oe7, 3oee, 3oeh,3ofn - yF1-ATPS (mutant)
    
  • 8f2k - yF1-ATPS + ATP + inhibitor – Cryo EM
    
  • 7tjs – yATPS + ADP + ATP - Cryo EM
    
  • 6cp3 – yATPS + oligomycin
    
  • 6cp5 – yATPS + oligomycin inhibitor – Cryo EM
    
  • 2wpd – yATPS + ADP + ATP
    
  • 7md3 – yATPS + ATP + drug – Cryo EM
    
  • 2wpd, 2wss - yF1-ATPS+subunit 9+Mg+ADP+ATP
    
  • 6tdu, 6tdv, 6tdw, 6tdx, 6tdy, 6tdz, 6te0 – ATPS + ADP + ATP – Euglena gracilis – Cryo EM
    
  • 5fl7 – ATPS + ADP + ATP – Yarrowiya lipolytica
    
  • 7jg5, 7jg6, 7jg7, 7jgb, 7jgc, 7nk7, 7nk9, 7nkb, 7nkd, 7nkh, 7nkj, 7nkk, 7nkl, 7nkn, 7nkp, 7nkq, 7nl9 – MsATPS – Cryo EM
    
  • 7jg8, 7jg9, 7jga – MsATPS + drug – Cryo EM
    
  • 7njk, 7njl, 7njm, 7njn, 7njo, 7njp, 7njq, 7njr, 7njs, 7njt, 7nju, 7njv, 7njw, 7njx, 7njy – MsATPS + ADP + ATP – Cryo EM
    
  • 6f36, 6rd4, 6rd5, 6rd6, 6rd7, 6rd8, 6rd9, 6rda, 6rdb, 6rdc, 6rdd, 6rde, 6rdf, 6rdg, 6rdi, 6rdj, 6rdk, 6rdl, 6rdm, 6rdn, 6rdp, 6rdq, 6rds, 6rdt, 6rdu, 6rdv, 6rdx, 6rdy, 6rdz, 6re0, 6re1, 6re2, 6re3, 6re4, 6re5, 6re6, 6re7, 6re8, 6re9, 6rea, 6reb, 6rec, 6red, 6ree, 6ref, 6rer, 6res, 6ret, 6reu – ATPS – Polytomella - Cryo EM
    
  • 5lqz, 5lqy, 5lqx – ATPS + ATPase inhibitor protein – Ogataea angusta - Cryo EM
    
  • 5dn6 – PdATPS + zeta inhibitor protein
    
  • 5t4o, 5t4p, 5t4q, 6oqr, 6oqs, 6oqt, 6oqu, 6oqv, 6oqw, 6pqv, 6vwk, 6wnq, 6wnr, 8dbv – EcATPS – Cryo EM
    
  • 7md2, 7tjt, 7tju, 7tjv, 7tjw, 7tjx – EcATPS + ATP – Cryo EM
    
  • 8dbp, 8dbq, 8dbr, 8dbs, 8dbt, 8dbu, 8dbw – EcATPS + ADP + ATP – Cryo EM
    
  • 3oaa – EcATPS (mutant) + ADP + ANP
    
  • 7xkh, 7xko, 7xkp, 7xkq, 7xkr – BaATPS – Cryo EM
    
  • 6n2d, 6n2y, 6n2z, 6n30 – BaATPS + ADP + ATP – Cryo EM
    
  • 7p2y, 7p3n, 7p3w – AbATPS + ADP + ATP – Cryo EM
    
  • 6tmg, 6tmh, 6tmi, 6tmj, 6tmk, 6tml – ATPS + ATPase inhibitor protein + OSCP + ADP + ATP – Toxoplasma gondii - Cryo EM
    
  • 6ynv, 6ynw, 6ynx, 6yny, 6ynz, 6yo0 – ATPS + ADP + ATP – Tetrahymena thermophila - Cryo EM
    
  • 8apa, 8apc, 8apd, 8ape, 8apg, 8apj, 8apk – TbATPS - Cryo EM
    
  • 8j0t – MtATPS - Cryo EM
    
  • 8jr0 – MtATPS + ATP + ADP + inhibitor - Cryo EM
    
  • 8j0s – MtATPS + drug - Cryo EM
    

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Michal Harel, Joel L. Sussman, Openfox